Jon A. Baskin - Texas A&M University-Kingsville

Some information on Texas marine reptiles which are not archosaurs is also presented.  Scroll down or click here.

LONE STAR DINOSAURS

The definitive and necessary reference: Lone Star Dinosaurs. by Louis Jacobs. College Station: Texas A& M University Press. 1995. 160p.

The Lone Star dinosaur exhibit sponsored by the Fort Worth Museum of Science and History and the Shuler Museum of Paleontology at Southern Methodist University is based on Dr. Louis Jacobs' book, Lone Star Dinosaurs (1995, Texas A&M University Press), featuring the original artwork of Karen Carr, whose official website is at: www.karencarr.com.

T. Michael Keesey's "The Dinosauricon" used to list Dinosaurs by Location, including those from Texas, but at present (8/2000) no longer does so.  However you can download (in WordPerfect format) a list of dinosaurs by continent.

Texas dinosaurs are also discussed and illustrated on Texas Parks and Wildlife / kids' page.  The dinosaurs of Texas lists them (note the differences with Keesey's page.)

A list of Texas dinosaurs from Dinodata also lists Texas dinosaurs by formation and provides links to further information.

The archosaurs from North-Central Texas are summarized in Jacobs and Winkler (1998).

Somewhat dated lists of Texas dinosaur distribution are in Weishampel (1990).

Lundelius (1986) includes a summary of Cretaceous fossils from the Balcones Fault trend.

Texas Dinosaurs: Life and Death in the Big Bend -- an exhibit from the Dallas Museum of Natural History

TEXAS DINOSAURS

Systematics

Superorder Dinosauria
     Order Saurischia
          Suborder THEROPODA
               Infraorder Herrerasauria
                    ? Herrerasauria incertae sedis
                         Protoavis texensis
                         Spinosuchus caseanus (may not be a dinosaur)
                    Family Herrerasauridae
                         Caseosaurus crosbyensis
               Infraorder Carnosauria
                    Superfamily Allosauroidea
                         Acrocanthosaurus atokensis
               Infraorder Coelurosauria
                    Superfamily Tyrannosauroidea
                         Family Tyrranosauridae
                              Tyrannosaurus rex
                         Saurornitholestes cf. langstoni
                    Maniraptora
                         cf. Deinonychus spp.
                         Ricardoestesia cf. gilmorei
                         Ricardoestesia isoceles
       Suborder Sauropodamorpha
               Infraorder SAUROPODA
                    ?Family Brachiosauridae
                         Paluxysaurus jonesi
                    Family Titanosauridae
                         Alamosaurus sanjuanensis

     Order Ornithischia
           Suborder FABROSAURIA
                 Family Fabrosauridae
                      Technosaurus smalli
                      Tecovasaurus murryi (may not be a dinosaur)
           Suborder ORNITHOPODA
                 Family Hypsilophodontidae
                      new species near  Hypsilophodon foxi
                 Family Iguanodontidae
                      Tenontosaurus dossi
                      Tenontosaurus tilletti
                 Family Hadrosauridae
                     Protohadros byrdii
                     Gryposaurus sp. (Kritosaurus cf. navajovius)
                     cf. Edmontosaurus sp.
                     lambeosaurine indet.
                     unnamed hadrosaur 

           Suborder Thyreophora
                 Infraorder ANKYLOSAURIA
                      Family Nodosauridae
                           Texasetes pleurohalio
                           Pawpawsaurus campbelli
                           Edmontonia rugosidens
                           unnamed ankylosaurs
           Suborder Marginocephalia
                 Infraorder PACHYCEPHALOSAURIA
                        Family Pachcephalosauridae
                             Stegoceras sp.
                 Infraorder CERATOPSIA
                        Family Ceratopsidae
                             Chasmosaurus mariscalensis
                             Torosaurus cf. latus
                             undetermined genus and species

Texas Dinosaurs by Time

 _____________________________________________________________________
|late LATE CRETACEOUS          Big Bend Region  -  West Texas         |
|_____________________________________________________________________|
|                  |        Black Peaks Formation                     |
|  Maastrichtian   |        Javelina Formation   El Picacho Formation |
|__________________|__________________________________________________|
|                  |         Aguja Formation     San Carlos Formation |
|   Campanian      |         Pen Formation                            |
|                  |                                                  |
|   Santonian      |                                                  |
|   Coniacian      |                                                  |
|__________________|__________________________________________________|
 

Tyrannosaurus rex?
Ricardoestesia
Saurornitholestes
Alamosaurus sanjuanensis
Gryposaurus sp.
cf. Edmontosaurus sp.
indeterminate lambeosaurine
Stegoceras sp.
Chasmosaurus mariscalensis
Torosaurus cf. latus
Edmontonia rugosidens
unnamed nodosaurids

 ______________________________________________
|early LATE CRETACEOUS    North-Central Texas  |
|______________________________________________|
|      Turonian           |                    |
|______________________________________________|
|    Cenomanian           | Woodbine Formation |
|______________________________________________|

Protohadros byrdii
unnamed nodosaurid
unnamed hadrosaur
cf. Ricardoestesia

 ____________________________________________________
|late EARLY CRETACEOUS       North-Central Texas     | 
|____________________________________________________|
|   |     |     |       |                            |
|_________|     |       |                            |
|   |     |     |  W    |                            |
|   |     |     |  A  G |                            |
|   |     |     |  C  r |    Paw Paw Formation       |
|   |     |     |  H  o |                            |
|   |     |     |  I  u |                            |
|   |     |     |  T  p |                            |
| E |  A  |  C  |  A    |                            |
| a |  L  |  O  |       |                            |
| r |  B  |  M  |       |                            | 
| l |  I  |  A  |_______|____________________________|     
| y |  A  |  N  |                                    |
|   |  N  |  C  |       Fredericksburg Group         |
| C |     |  H  |                                    |
| R |     |  E  |____________________________________|
| E |     |  E  | T    |                  |  A       |
| T |     |     | r  G | Paluxy Fm        |  n       |
| A |     |  S  | i  r |                  |  t  F    |
| C |_____|  E  | n  o | Glen Rose Fm     |  l  m    |
| E |  A  |  R  | i  u |                  |  e       |
| O |  P  |  I  | t  p | Twin Mountains Fm|  r       |
| U |  T  |  E  | y    |                  |  s       |
| S |  I  |  S  |      |                             |
|   |  A  |     |      |                             |
|   |  N  |     |      |                             |
|___|_____|_____|______|_____________________________|
 

Acrocanthosaurus atokensis
cf. Deinonychus
Paluxysaurus jonesi
new species near Hypsilophodon foxi
Tenontosaurus dossi
Tenontosaurus tilletti
Texasetes pleurohalio
Pawpawsaurus campbelli
unnamed nodosaurids

 _______________________________________________________________
|LATE TRIASSIC              Texas Panhandle, West Texas         |
|_______________________________________________________________|
|                  |                                            |
|    Rhaetian      |                                            |
|__________________|                                            |
|                  |   Dockum Group                             |
|    Norian        |                     Bull Canyon Member     |
|__________________|                                            |
|                  |                     Tecovas Member         |
|    Carnian       |                     Colorado City Member   |
|                  |                     Camp Springs Member    |
|__________________|____________________________________________|
 

Caseosaurus crosbyensis (may not be a dinosaur, but is likely a dinosauriform)
Protoavis texensis (in part)
Spinosuchus caseanus (may not be a dinosaur)
Technosaurus smalli (may not be a dinosaur)
prosauropod indet. (may not be a dinosaur)
Tecovasaurus murryi (may not be a dinosaur)

Nesbitt and others (2007) reviewed the putative record of late Triassic record dinosaurs in North America.  The only taxa from Texas that they assigned to the Dinosauria is Protavis texensis (in part).

DINOSAURIA

Order SAURISCHIA

Suborder THEROPODA

Classification

• Order Saurischia
  • Suborder Theropoda

Infraorder HERRERASAURIA

Classification

• Order Saurischia
  • Suborder Theropoda
    • Infraorder Herrerasauria
      • Family Herrerasauridae
        • Caseosaurus crosbyensis
        • probably Protoavis texensis in part
        • possibly Spinosuchus caseanus

Caseosaurus Hunt, Lucas, Heckert & Lockley, 1998

Type and only species: Caseosaurus crosbyensis Hunt, Lucas, Heckert, Sullivan & Lockley, 1998

Locality

• Tecovas Member, Dockum Formation, Crosby County, Texas;  latest Carnian (Latest Triassic)

Comment

• Based on a right ilium originally attributed to Chindesaurus bryansmalli (Long and Murry, 1995).    Nesbitt and others (2007) refer it to the Dinosauriformes and state that it cannot be unambiguously assigned to the Herrerasauridae or Dinosauria.

Protoavis Chatterjee, 1991

Type and only species: Protoavis texensis Chatterjee, 1991

:Locality

• Dockum Formation, Texas;  Norian (Late Triassic)

Comment

• This small (mass = 350 g) dinosaur (in part) is based on skeletal material from at least 2 individuals (possibly from different taxa).  Protoavis was originally described as the oldest bird (Chatterjee, 1986, 1991), significantly older than the late Jurassic Archaeopteryx, usually regarded as the ancestral bird. Further analysis indicated that certain parts may belong to a primitive theropod, others to a pterosaur.   Hunt and others (1998) conclude that it is probably an aberrant theropod.  Nesbitt and others (2007) refer the dinosaurian parts to Coelophysoidea indet.
  

Spinosuchus von Huene, 1932

Type and only species: S. caseanus von Huene, 1932

Locality

• Dockum Group, north of Cedar Mountain, Crosby County,  latest Carnian (Late Triassic)

Comment

• specimens consist of vertebrae.  Once thought to be a "pseudosuchian", but may be a theropod.  Murry (1986) considers that it is probably a rauisuchian or poposaurian.  Nesbitt and others (2007) refer it to Archosauriformes incertae sedis.

Infraorder CARNOSAURIA

Classification

• Order Saurischia
  • Suborder Theropoda
    • Infraorder Carnosauria
      • Family Allosauridae

Acrocanthosaurus

Type and only species: A. atokensis  Stovall and Langston, 1950

Classification

• Order Saurischia
  • Suborder Theropoda
    • Superfamily Allosauridoidea

Distribution

• Trinity Group of Texas; also Antlers Formation, Oklahoma.  Aptian-Albian; late Early Cretaceous.

Etc.

• The most complete skeleton known from Texas comes from the Twin Mountains Formation at Parker Ranch in Parker County (Harris, 1998).  Large isolated teeth attributable to this genus occur elsewhere in the Trinity.
• Check out Karen Carr's image of Acrocanthosaurus attacking Pleurocoelus for the Fort Worth Museum of Science and History.

© Karen Carr, used by permission of Louis L. Jacobs; click to enlarge

• The carnivorous Acrocanthosaurus was a large allosauroid, 8 to 12 meters long and weighing 2 to 3 tons.  Its name comes from the long vertebral spines along its back.  These have been interpreted as forming a sail.  However, Bailey (1997) interprets that these neural spines instead formed a broad (buffalo-like) hump.  See below for information on  footprints.
• Acrocanthosaurus is most closely related to the early Cretaceous, African genus Carcharodontosaurus.

Infraorder COELUROSAURIA

Classification

• Order Saurischia
  • Suborder Theropoda
    • Infraorder Coelurosauria
      • Tyransosauroidea
      • Maniraptora

Tyrannosaurus Osborn, 1905
Classification

• Order Saurischia
  • Suborder Theropoda
    • Infraorder Coelurosauria
      • Superfamily Tyrannosauroidea

Tyrannosaurus rex Osborn, 1905 (type)

Texas Locality

• Brewster County, lower part of the Tornillo Group; Maastrichtian (latest Cretaceous).

Texas material.

• This genus is recognized from Big Bend National Park from a maxilla with teeth described by Lawson (1976).  It was suggested that the relatively small size of this specimen may be because it is a young adult or because it is from a smaller theropod such as Albertosaurus. 
• Carr and Williamson (2000, 2004) identify the Tornillo Formation tyrannosaurid as T. rex.
•  Tyrannosaurid  teeth are also known from the Campanian Aguja formation (Sankey, 2001)
• Karen Carr's Big Bend T. rex.

Saurornitholestes Sues, 1970
Classification

• Order Saurischia
  • Suborder Theropoda
    • Infraorder Coelurosauria
      • Superfamily Tyrannosauroidea

Sauronitholestes cf. langstoni Sues, 1970

Texas Locality

• Aguja Formation, Campanian (late Cretaceous).

Texas material.

• teeth or teeth fragments (Sankey, 2001)

cf. Deinonychus

Classification

• Order Saurischia
  • Suborder Theropoda
    • Maniraptora

Texas Localities

• This genus has been reported from isolated teeth from the Trinity Group of Texas (Winkler and others, 1988; Jacobs, 1995), but the material has not been described in detail .  Brinkman and others (1998) note that the dromaeosaurid teeth from the lower Trinity Group are considerably smaller than those from the Cloverly Formation (type locality) of Montana and Wyoming and those from the upper Trinity are the same size or larger then the Cloverly Deinonychus.
• Associations of Tenontosaurus and its predator Deinonychus have been discovered in Montana and Oklahoma, and possibly in Texas (a few dromaeosaurid teeth occur with the Doss Ranch specimens).

Ricardoestesia Currie, Rigby, and Sloan, 1990

Classification

• Order Saurischia
  • Suborder Theropoda
    • Infraorder Maniraptora
      • family unknown

Ricardoestia cf. gilmorei Currie, Rigby, and Sloan, 1990
Ricardoestia isosceles Sankey, 2001

Texas Localities

• This genus is tentatively recognized from the Cenomanian Woodbine Formation (Lee, 1997).   Ricardoestia cf. gilmorei and R. isosceles occur in the Judithian (Campanian) Aguja Formation (Sankey, 2001).

Theropoda indet.

• Isolated teeth from the Aguja Formation referred to the maniraptoran cf. Dromaeosaurus (Rowe and others, 1992) have been reassigned to theropod genus and species undetermined (Sankey, 2001).

Suborder SAUROPODA

Classification

• Order Saurischia
  • Suborder Sauropodamorpha
    • Infraorder Sauropoda

Paluxysaurus

Classification

• Order Sauropodamorpha
  • Infraorder Sauropoda
    • ?Family Brachiosauridae

Type and only species: Paluxysaurus jonesi Rose 2007

Locations

• Glen Rose Formation and Paluxy Formation; early Albian (about 110 Ma), late early Cretaceous.

Comments

• Rose (2007) established a new genus and species for the early Cretaceous sauropod from Texas. 
• This dinosaur made the huge "wash basin" footprints in the early Cretaceous, Glen Rose Formation.

            Image from Roland T. Bird’s, “A Dinosaur Walks into the Museum,” published in Natural History, February 1941., reprinted with permission.

• Previously it was referred to Pleurocoelus, a brontosaur seventy feet long and weighing twenty tons.  However,  assignment to this genus was considered uncertain (Langston, 1974).   Pleurocoelus is sometimes considered a junior synonym of Astrodon (e.g., Brinkman, Cifelli, and Czaplewski, 1998), the state fossil of Maryland.   Wedel, Cifelli, and Sanders (2000) described Sauroposeidon proteles, a new brachiosaurid sauropod from the early Cretaceous Antlers Formation of Oklahoma and suggest that other material referred to Pleurocoelus needed to be reexamined.  Rose (2007) distinguished the new Texas genus from previously named taxa, and concludes that both Pleurocoelus and Astrodon should be considered incertae sedis.
• Karen Carr's image for Lone Star Dinosaurs shows that even a Paluxysaurus could become a meal for Acrocanthosaurus.

© Karen Carr, used by permission of Louis L. Jacobs; click to enlarge

• The state dinosaur of Texas.  

Tidwell and Carpenter (2003) describe the braincase of a titanosauriform sauropod from the Glen Rose Formation in Blanco County.  It is described as having strong similarilarities with brachiosaurids and titanosaurids.

Alamosaurus

Classification

• Order Saurischia
  • Suborder Sauropoda
    • Family Titanosauridae

Distribution

• Texas, Big Bend: Black Peaks, Javelina, and El Picacho Formations; also New Mexico, Utah. Maastrichtian (latest Cretaceous)
• Preliminary report on a new sauropod locality in the Javelina Formation at Big Bend

Comments

• Alamosaurus was 21 m long and weighed 30 tons.  It is the only North American sauropod from the Late Cretaceous (Maastrichtian).  Although it is known from Texas, it is not named after the Alamo in San Antonio, but after the Ojo Alamo Formation (the type locality is now recognized as the Kirtland Shale), at the Ojo Alamo (Spanish for cottonwood spring) trading post in the San Juan Basin, New Mexico.
• Alamosaurus may represent an invasion from South America by titanosaurids, following the mid-Cretaceous extinction of the brachiosaurids in North America.   Lehman and Coulson (2002) conclude that Alamosaurus is most closely related to titanosaurs from Brazil and Argentina.

DINOSAURIA

Order ORNITHISCHIA

Suborder FABROSAURIA

Classification

• Order Ornithischia
  • Suborder Fabrosauria
    • Family Fabrosauridae
      • Technosaurus smalli
      • Tecovasaurus murryi

Technosaurus Chatterjee, 1984.

Type and only species:  T. smalli Chatterjee, 1984 (type)

Location and age: Dockum Group, Texas Panhandle, Norian (late Triassic).

Comment:

• This 1 to 2 m long putative primitive dinosaur is known only from a right lower jaw.   A premaxilla and a posterior lower jaw that had been assigned to this taxon were reassigned to an indeterminate prosauropod (Sereno, 1992; Hunt and others, 1998).  Nesbitt and others (2007) consider the specimens possible Dinosauromorpha
  

Tecovasaurus Hunt & Lucas, 1994

Type and only species: T. murryi Hunt & Lucas, 1994 (type)

Localities

• Tecovas Formation, Crosby County, Texas and New Mexico and Arizona; Carnian (late Triassic)

Comment

• This taxon is based on isolated teeth.  Irmis (2005) notes that isolated teeth cannot be used to assign specimens to the Ornithischia and instead refers Tecovasaurus to Archosauriformes incetae sedis.

Suborder ORNITHOPODA

Classification

• Order Ornithischia
  • Suborder Ornithopoda

Family HYPSILOPHODONTIDAE

Hypsilophodont - new species, similar to Hypsilophodon foxi (see Winkler and others, 1988; Winkler and Murry, 1989)

Classification

• Order Ornithischia
  • Suborder Ornithopoda
    • Family Hypsilophodontidae

Location

• Twin Mountains Formation, Proctor Lake, Commanche County, Texas, late Aptian (late Early Cretaceous), about 115 Ma.

Comments

• Karen Carr's image for Lone Star Dinosaurs shows a group of  these small hypsilophodonts cavorting in front of a Tenontosaurus.   This temporary display at the Dallas Museum of Natural History shows a skeleton of this as-yet, unnamed primitive ornithopod.

© Karen Carr, used by permission of Louis L. Jacobs; click to enlarge

• The Proctor Lake locality, discovered by Tarleton State University student Rusty Branch, has a few adults and clusters of juvenile hypsilophodontids, possibly indicating a nesting site, although no eggs have been found. (Winkler and Murry, 1989).

IGUANODONTIA

Tenontosaurus

Classification

• Order Ornithischia
  • Suborder Ornithopoda
    • Family Iguanodontidae

Comments

• Tenontosaurus is the large dinosaur in the illustration by Karen Carr  for Lone Star Dinosaurs 

© Karen Carr, used by permission of Louis L. Jacobs; click to enlarge

• Tenontosaurus is the most common large herbivore known from the late early Cretaceous of North America. 
• Tenontosaurus is intermediate in morphology between hypsilophodontids and iguanodonts (Winkler and others, 1997).
• Associations of Tenontosaurus and its predator Deinonychus have been discovered in Montana and Oklahoma, and possibly in Texas (a few dromaeosaurid teeth occur with the Doss Ranch specimens).

Tenontosaurus dossi Winkler and others, 1997.

Location

• Twin Mountains Formation, Parker County, Texas, latest Aptian (approximately 113 Ma)

Comments

• In 1988, seven-year-old Thad Williams and his father Ted discovered this new species of primitive iguanodont.
• Tenontosaurus dossi is slightly more primitive than Tenontosaurus tilletti.

Tenontosaurus tilletti?

Comment

• Tenontosaurus tilletti is known from the Cloverly Formation, Montana. This species probably occurs in higher levels of the Trinity Group (Glen Rose and Paluxy Formations) of Texas (Winkler and others, 1997).

Family HADROSAURIDAE

Subfamily Hadrosaurinae

Classification

• Order Ornithischia
• Suborder Ornithopoda
  • Family Hadrosauridae
    • Subfamily Hadrosaurinae
      • Protohadros byrdii
      • unnamed hadrosaur
      • Kritosaurus navajovius

Protohadros Head, 1998

Type and only species: Protohadros byrdii Head 1998

Location

• Flower Mound (north of Dallas-Fort Worth, Texas), Woodbine Formation, Cenomanian (early late Cretaceous), 95 Ma.

Comment

• Protohadros is intermediate in morphology between the iguanodonts and the other hadrosaurids (Head, 1998).  It is the one of the oldest and most primitive named hadrosaurids, a group that was to become increasingly important in the late Cretaceous.

Main and Fiorello (2003) report finding hadrosaur postcrania along the shores of Lake Grapevine from the Woodbine Formation.

Kritosaurus Brown, 1910 or Gryposaurus Lambe, 1914

Texas species:

• Kritosaurus cf. navajovius (or n.sp. or ?Gryposaurus n. sp). from the Pen Formation, San Carlos Formation, Aguja Formation, and El Picacho Formation. 

Comment

• Because Kritosaurus navajovius (Brown, 1910) is based on very poor material,   Gryposaurus is the preferred name for this hadrosaurine according to Weishampel and Horner (1990).  However, Rowe and others (1992) refer some of the Aguja hadrosaurines to Kritosaurus.  However, reversing their opinion of the previous year Hunt and Lucas (1993) conclude that Kritosaurus navajovius is a nomen dubium.  Sankey (2001) stated that Kritosaurus cf. navajovius and an unidentified labeosaurine and an indeterminate hadrosaurid occur in the Aguja Formation.  In an abstract, Wagner and Lehmann (2001) refer the Aguja hadrosaur toKritosaurus n.sp.  Gates and Sampson (2007) suggest the Aguja material may be attributable to Gryposaurus n. sp.

Suborder Marginocephalia

Infraorder PACHYCEPHALOSAURIA
Family Pachcephalosauridae

Stegoceras sp. has been reported from the San Carlos Formation (Lehmann cited in Weishampel, 1990).  Teeth from the Aguja Formation assigned to cf. Troodon (Rowe et al., 1992) have been identified as belonging to a pachycephalosaurid.
         

Infraorder CERATOPSIA

Classification

• Order Ornithischia
  • Suborder Marginocephalia
    • Infraorder Ceratopsia
      • Family Ceratopsidae
        • Subfamily Chasmosaurinae
          • Agujaceratops mariscalensis
          • Torosaurus utahensis
          • undetermined genus and species

Comment

• In Texas, ceratopsian dinosaurs occur in Campanian and Maastrichtian (late Cretaceous) sediments in the Big Bend area.

Agujaceratops

Chasmosaurus mariscalensis Lehman, 1989
Agujaceratops mariscalensis Lucas, Sullivan, and Hunt 2006

Distribution

• Aguja Formation, Texas;  Campanian (Judithian) (late Cretaceous)

Comments

• A nearly complete skull of this ceratopsian is described by Forster and others (1993).  Lucas, Sullivan, and Hunt (2006) established a new genus Agujaceratops for this species.

Torosaurus Marsh, 1891

Type species

• Torosaurus latus Marsh, 1891

Distribution

• late Maastrichtian (Lancian; Late Cretaceous).  Torosaurus, originally described from Wyoming, is also known from the Javelina Formation, Tornillo member, Texas, and elsewhere.  Lawson (1976) referred two Texas specimens to Torosaurus utahensis.  Farke (2002) stated that one of these specimens, a partial right parietal, can be referred to Torosaurus sp.; the other, a partial orbital horn, to Chasmosaurinae indet.  Sulivan, Boere, and Lucas (2005) consider the Big Bend specimens (Lawson, 1976; Lehman, 1996) as indeterminate chasmosaurines.

Comments

• This ceratopsian had one of the largest skulls of any terrestrial vertebrate, approximately 8 feet long (exceeded only by a specimen of Pentaceratops (Lehman, 1998).
• Size: 6-8 m (no complete skeletons are known), 7-8 tons.

Genus and species undetermined Lehman 1996

Distribution

• Picacho Formation, Texas;  Maastrichtian (latest Cretaceous)

Suborder Thyreophora

Infraorder ANKYLOSAURIA

Classification

• Order Ornithischia
  • Suborder Thyreophora
    • Infraorder Ankylosauria
      • Family Nodosauridae

Comment

• Texas nodosaurids (such as Pawpawsaurus) are found in early and late Cretaceous marine sediments in Texas and elsewhere.  They frequented the coast of the ancient sea.

Pawpawsaurus Lee, 1996

Type and only species: Pawpawsaurus campbelli Lee, 1996

Location

• Paw Paw Formation, Tarrant County (north of Fort Worth), Texas, late Albian (latest Early Cretaceous), about 100 Ma.

Material

• A skull, with bony eyelids (Lee, 1996).  Cameron Campbell has a  website devoted just to his Texas Pawpawsaurus .

Texasetes Coombs, 1995

Type and only species: Texasetes pleurohalio Coombs, 1995.

Location

• Paw Paw Formation, Tarrant County (north of Fort Worth), Texas, late Albian (latest Early Cretaceous), about 100 Ma.

Material

• two teeth and much of a skeleton (Coombs, 1995) of a nodosaur about 3 m long.

Comment

• According to Lee (1996), this material is not diagnostic and can not be demonstrated to be the same taxon as Pawpawsaurus.

Nodosauridae indet.

• Unnamed nodosaurid material occurs in the Woodbine Formation, and includes a skeleton of a baby (Jacobs, 1995; Lee, 1997).

Edmontonia Sternberg, 1928

Texas species: Edmontonia rugosidens (Gilmore, 1930) is reported from the Aguja formation.

TRACE FOSSILS

THE GLEN ROSE TRACKWAY

• Hawthorne (1990) provides a good introduction to Glen Rose trackways.
• The large dinosaur footprints in the Glen Rose trackways have been assigned to the ichnotaxon Brontopodus birdi (Farlow and others, 1989),  named in honor of Roland T. Bird, who excavated a long trackway for the American Museum of Natural History.  They were likely made by Paluxysaurus (formerly Pleurocoelus). Wedel and Cifelli (2006) state that the tracks are so large they must have been made by the giant sauropod Sauroposeidon. Theropod trackways have been referred to Grallator sp. and were probably made by Acrocanthosaurus. (Rogers, 2002).
• Dinosaur Valley State Park in the early Cretaceous, Glen Rose Formation.
• Learn more about the Paluxy River tracksite and more from Glen Kuban.  There are no human footprints here (Milne and Schafersman, 1983).
• There are numerous other tracksites in the Glen Rose Formation, such as those at Davenport Ranch (Lockley and Hunt, 1995), Kinney County (Rogers, 2002) and Boerne (Hawthorne et al., 2002).

Dinosaur footprint in Glen Rose Formation near Tarpley, Texas. Click on image to enlarge

Other Texas Mesozoic Life

Other Texas Archosaurs

Order Thecodontia

Suborder PSEUDOSUCHIA

Shuvosaurus

Type and only species

• S. inexpectatus Chatterjee, 1993

Location

• Post Quarry, Garza County, Bull Canyon Formation, Dockum Group, Norian, late Triassic

Comment

• Chatterjee (1993) described this taxon, based on skull fragments, an edentulous dentary, and a dorsal vertebra, as an ornithomimosaurian theropod.  Long and Murry (1995) suggest that it may be a ?rauisuchian that they name Chatterjeea elegans.   Hunt and others (1998) note that Shuvosaurus lacks any dinosaurian synapomorphies and tentatively support Long and Murry's interpretation. Rauhut (1997) suggested it is an early coelophysid.  Nesbitt and Norell (2006) conclude that Shuvosaurus is related to the early suchian crocodylian Effigia, a toothless form convergent with the ornithomimids.

Suborder Aetosauria

Family Steganolepididae

• Coahomasuchus kahleorum

  • Howard County, Texas, Colorado City Formation, Dockum Group, late Carnian, late Triassic (Heckert and Lucas, 1999)

• Desmatosuchus smalli  (Small, 1989; 2002; Parker, 2005)
• Tecovasuchus chatterjeei (Martz and Small, 2006)
Paratypothorax sp.
Typothorax coccinarum
• Post (Miller) Quarry, Garza County, Bull Canyon Formation, Dockum Group, Norian, late Triassic (Small, 1989; 2002; Parker, 2005)

Suborder Phytosauria

Family Phytosauridae

• Rutiodon
• Angistorhinus
• Paleorhinus

Triassic phytosaurs resemble modern alligators which inhabit southeast Texas today, but the resemblance is only superficial. For example, the nostrils of an alligator are on the tip of the snout, while the nostrils of a phytosaur are between its eyes. The similarity of appearance results from convergent evolution - adaptation to a similar kind of lifestyle by different kinds of animals.

RAUISUCHIA

Postosuchus kirkpatricki
Chatterjeea elegans

Location

• Post Quarry, Garza County, Bull Canyon Formation, Dockum Group, Norian, late Triassic (Parker, 2005), Tecovas and Copper Canyon Formations, Dockum Group, Norian, late Triassic

Order CROCODYLIA

Pachycheilosuchus trinquei

Glen Rose Formation

• Rogers (2003) described this mesoeucrocodylian from near-shore, marine-to-brackish deposits of the Albian Glen Rose Formation in Erath County.   He notes the presence of  other Glen Rose crocodylians, particularly the "Glen Rose Form" (Langston, 1974).  Estimated body length is 60-80 cm.  Rogers (2000) has also described an egg from this species.

Deinosuchus rugosus

(=Phobosuchus riograndensis)

upper Aguja Formation

• Known (until recently) from two skulls, each about 2 m long.   One of the largest extinct crocodilian yet found, Deinosuchus may have preyed on smaller dinosaurs.  Deinosuchus had been estimated to be over 45 feet in length extrapolated from skull size (largest recent crocodile is around 33 ft).  Recent discoveries indicate that it was 9 to 12 meters long and had a body mass of 2,500 to 5,000 kg, three to five times more than the largest crocodiles alive today (Erickson and Brochu, 1999; Schwimmer, 2002).
• Deinosuchus from the Dallas Museum of Natural History
• Which was bigger (by D. L. Parsell, National Geographic News) Deinosuchus or Sarcosuchus (supercroc)

(crocodile classification: Brochu, 1997, Jour. Vert. Paleont., 17:679-697; 2003, Ann. Rev. Earth Planet. Sci., 31:357-397.)

Goniopholus cf. G. kirtlandicus

• Aguja formation

Woodbineosuchus byersmaurecei Lee, 1997

• Woodbine Formation.  A narrow snouted, fish-eating goniopholid.
  

Order Pterosauria

Pterosaurs soared over the oceans of Texas more than 100 million years ago

Eudimorphodon

• Dockum Group, Carnian (220 million years ago)
• This record (Murry, 1986) may be the earliest for pterosaurs (Andres, 2006)

Quetzalcoatlus northropi

• The great Texas pterosaur
• Quetzalcoatlus was the largest animal ever to fly under its own power, having a wing span of 11-12 m long (almost forty feet) and a weight of about 90 kg..

Quetzalcoatlus sp. Keller and Langston, 1996, JVP 16:222-231.

Coloborhynchus wadleighi Lee, 1994

• Paw Paw Formation

Azhdarichiidae indet.

• Glen Rose Fm.

Pterosauria indet.

• Antlers Fm.

Marine Reptiles

PLESIOSAURS

These aquatic sauropterygian lepidosauromorphs of the Mesozoic epicontinental seaways have four paddle-shaped (Wing-fins of Adams, 1998) limbs.  Both long necked (plesiosauroids or elasmosaurs) and short necked forms (pliosauroids) are present in Texas.   In a published abstract of his unpublshed 1981 thesis, Storrs (1983) states that nine genera of plesiosaurs are known from Texas.  The abstract briefly discusses four genera: Trinacromerum, Thalassomedon, ?Alzadasaurus, and Polyptychodon.

Libonectes morgani (Welles 1949) is a very long-necked elasmosaurid plesiosaur from the Britton Formation, Eagle Ford Group, near Cedar Hill, Dallas County, Texas.

Ceraunosaurus brownorum Thurmond is a short-necked pliosaur from the Lake Waco Formation (Cenomanian).  Storrs (1981) states that this genus "is an obvious specific variant of Tricanomerum."

Brachauchenius lucasi, a very large pliosaur, was first collected in   Kansas (Williston, 1903). A second specimen (UNSM 2361) was collected from the Eagle Ford formation near Austin, Texas (Williston, 1907).  A skull from Kansas is about  1.5 m. long.

Trinacromerum bonneri Adams is known from the latest Cretaceous Taylor Marl, as well as the type locality in South Dakota.  Trinacromerum is a pliosaur, all high speed swimmers, of which this is the fastest.

MOSASAURS

These are giant marine, varanoid lizards, that may be ancestral to snakes.  Their diet included ammonites.   Recently, Tomoki Kase (Geology, 1998) indicated that the putative puncture marks are weakened areas of the shell where limpets attached to the dead ammonites' shells.  This has been disputed and evidence for mosasaur predation on ammonites has been strongly supported  (Tsujita and Westermann, Palaeogeography, Palaeoclimatology, Palaeoecology, 2001; Kauffman, PALAIOS, 2004).

Tylosaurus proriger.

• This 10 meter-long, late Cretaceous mosasaur is on display at the Dallas Museum of Natural History.

Mosasaurus maximus: the Onion Creek mosasaur

Halisaurus

• Caldwell and Bell refer a series of dorsal vertebrae from the Upper Cretaceous Pen Formation to this genus

Globidens

• Polcyn and Bell (2005a) recognize this rare durophagous mososaur fron North-Central Texas.  The note that Platecarpus intermedius belongs in this genus.

Dallasaurus turneri

• Bell and Polcyn (2005) describe a primitive mososaur with four limbs that was discovered near Dallas.

Russellosaurus coheni

• This new genus is described by Polcyn and Bell (2005b) from Cedar Hill, Dallas County, Texas.  This 92 Ma species is represented by the oldest well-preserved mosasaur skull from North America.

ICTHYOSAURS

Ichthyosaurs are the most fully-aquatic reptiles.  They are now classified as primitive diapsids

Main and Fiorello (2002) note in an abstract that the common Cretaceous genus Platypterygius is known from the Grayson Marl (lower Cretaceous) in Tarrant County.  Platypterygius has a better fossil record in Australia

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